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  • Article title
  • Abstract
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  • Introduction
  • Materials and methods
    • Ethical statements
    • Specimen sampling
    • Morphological study
    • Molecular study and phylogenetic analyses
    • Morphological abbreviations
  • Results
    • Systematics
    • Molecular phylogenetic analyses
  • Discussion
  • Acknowledgements
  • References
  • Supplementary material
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