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Article title
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Abstract
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Key Words
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Introduction
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Materials and methods
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Specimens
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Morphological data
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DNA extraction, amplification and sequencing
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Phylogenetic analyses
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Divergence-time estimation
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Taxonomic accounts
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Results
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Phylogenetic analyses and divergence dating
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Taxonomical accounts
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Discussion
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Phylogenetic relationships
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Timing of species diversification and biogeographical implications
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Acknowledgements
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References
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Supplementary material
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