Important characters for the distinction of the Calumma furcifer group (n = 12; blue diamonds) and the C. nasutum group (n = 12; green squares), including the assignment of C. vatosoa (n = 4, red triangles) and C. peyrierasi (n = 4; yellow dots). Abbreviations: NSA, number of scales on lower arm in a line from elbow to manus; NIL, number of infralabial scales; DSA, diameter of largest scale on lower arm; ROSSC, ratio of distance from the anterior margin of the orbit to the snout tip and snout-casque length.