Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16S and ATPα (Suppl. material 1: Table S1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the “Systematic checklist” for more information). Small green (PP ≥ 0.95) and red (PP < 0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1–32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods).