Corresponding author: Mark-Oliver Rödel (
Academic editor: Johannes Penner
Herein, we describe the tadpoles of six
The anuran family
During the last 15 years we collected numerous tadpoles of morphologically similar, torrenticolous arthroleptid tadpoles, apparently belonging to 10 species of
Tadpoles were collected with dip nets and anesthetized in chlorobutanol solutions before being either preserved in formalin (5–10%) or ethanol (75%). All were finally stored in 75% ethanol. Prior to preservation tail tips of representative specimens were preserved separately in 96% ethanol for genetic investigations. All tadpole vouchers and tissue samples are deposited at the collection of the
All measurements were taken by FG and MR. Measurements of randomly chosen vouchers were measured by both persons and the entire dataset only put together after these measures did not differ. Measures were taken with a digital caliper (accuracy ± 0.1 mm) and a measuring ocular on a Leica MZ95 dissecting microscope (accuracy ± 0.02 mm).
The following measurements were taken:
In the text, measurements or ratios are usually summarized (for
Photos of entire tadpoles were taken with a Canon EOS 50D digital camera and a 50 mm 1:2.5 lens. Mouthpart pictures were taken with a stacking camera (Leica DFC490) on a Leica Z16 APO A microscope. Single exposures were combined using the Automontage® software v. 5.03.0061 (Syncroscopy). All pictures were edited with Adobe Photoshop v. 18.1.0. Schematic sketches of the keratodont formulae are usually based on several individuals of the respective species.
To test, if and how the tadpoles of the different species can be distinguishable, we plotted indices of our various measures against each other and ran a Principle Component Analysis (PCA) using qti-Plot and Past 3.18 (
All tadpoles were caught in small to medium-sized streams. The description is mainly based on
Long slender tadpole with long, muscular tail and narrow fins (Fig.
The largest tadpole, still Gosner stage 25 (
Brown, slightly fading from snout to tail; dorsal surfaces with irregular dark speckles; ventral surfaces without or only few darker markings; tail fins sometimes with dark blotches or speckles, partly transparent. Life color of very small specimens with much yolk, were much lighter than larger specimens (Fig.
Young
According to
Whereas all genotyped tadpoles were genetically almost identical amongst each other and an adult from Mount Kala, Cameroon, further genetic comparisons with specimens from across the range of the species revealed three distinct genetic lineages (results not shown). Our tadpoles originated from Cameroon. All genetic samples from Gabon and the Republic of Congo, fell into a different clade. As the type locality of
The two specimens from Mount Manengouba were caught in medium-sized streams in gallery forest; the Korup specimens were from small creeks in rainforest. The description is mainly based on
Long robust tadpole with muscular tail (Fig.
The largest tadpole (
Brown to dark brown, dorsal surfaces darker, venter lighter and more grayish; flanks densely beset with diffuse and small brown spots; intensity of patterning deceases from back to vent; tail fins slightly transparent, dorsal fin with dark patterning, ventral fin with fewer dark spots.
All tadpoles were caught in forest streams. The description is mainly based on
Long slender tadpole with long, muscular tail (Fig.
The tadpoles (all Gosner stage 25) ranged from 45 mm to 74 mm total length.
Basic color light brown, the dorsal surfaces darker; back and tail with irregular, big, dark brown spots or blotches; venter light with few or no dark speckling; tail fins beige, semitransparent with dark brown spots.
Morphologically and genetically tadpoles of this species were very similar to
The tadpole was caught in a forest stream. The genotyped tadpole has been compared with the two adults from Mount Nlonako, Cameroon (
Long slender tadpole with long, muscular tail (Fig.
The tadpole (Gosner stage 25) measures 97.4 mm in total length.
Basic color light brown with darker and speckled dorsal surface; venter light and grayish with few or no dark speckling; tail fins beige, semitransparent with irregular tiny dark brown spots. In life the color was of a duller, darker brown (Fig.
The tadpole was caught in a forest stream. It was genotyped and compared with an adult from east of Ntale village, Banyang-Mbo, Cameroon (MCZ A-136785; GenBank
Long slender tadpole with long, muscular tail (Fig.
The tadpole (Gosner stage 25) measures 51.7 mm in total length.
Basic color light brown, the dorsal surfaces darker; back and tail axis with irregular, dark brown spotted patterns; ventral surfaces light with few or no dark speckling; tail fins beige, semitransparent with tiny dark brown spots on dorsal fin.
Long tadpole with muscular tail; body oval in dorsal and lateral view (Fig.
The maximum size of
General color light brown or beige, with irregular small brown spots covering body and tail; ventral surfaces of body more grayish, dark speckling almost absent; tail tip darker.
All tadpoles were caught in small to medium-sized streams. The description is mainly based on
Robust tadpole with long muscular tail (Fig.
The largest tadpole in Gosner stage 25 has a body length of 22.5 mm; the longest one (Gosner stage 28) had a total length of 73.2 mm.
Basic color brownish, dorsal surfaces darker than ventral ones, turning into yellow, tail lighter brown; tail axis with few dark speckles; tail fins very light brown, semitransparent and with a longitudinal row of small spots on the margin of dorsal fin.
As in
The description is mainly based on the two tadpoles accessioned as
Long slender tadpole with strong, muscular tail (Fig.
The largest tadpole in Gosner stage 25 measured 95.9 mm total length. Gosner stage 41 tadpole measured 77.5 mm. Tadpoles with more than 10 cm total length have been caught (Guibé and Lamotte 1958b; MOR unpubl. data).
Body more or less uniform dark brown; ventral lighter, slightly grayish; tail fin margins are missing dark pigmentation; last third of tail darker to almost black (Fig.
Life coloration of an
All tadpoles were caught in small to medium-sized streams and rivers in secondary and very degraded forests. The description is mainly based on
Long slender tadpole with strong and muscular tail (Fig.
The largest tadpole in Gosner stage 25 measured 26.2 mm body length and had an approximate total length of 78 mm. The most advanced specimen at Gosner stage 28 had a total length of 108.1 mm.
Body dark brown; tail very light brown to beige; lateral sacs and ventrum lighter brown or dark gray; a broad dark brown longitudinal band on tail axis reaching about middle to two-thirds of tail length, proximate end of tail with smaller dark speckles; fins light yellowish beige (Fig.
Life coloration of an
In larger specimens (Fig.
Whereas all genotyped tadpoles and the comparative adult were genetically almost identical, a single specimen (
The tadpole was caught in a medium-sized river within a heavily degraded forest fragment. The tail tip had been taken as tissue sample; the continuation of the tail tip was thus reconstructed based on the generalized tail shape of the genus, and total length estimated (compare Fig.
The tadpole’s sequence has been compared with an adult from Kodmin, Cameroon (
Long slender tadpole with narrow, muscular tail (Fig.
Total length of this tadpole was estimated to be about 18 mm (Fig.
Body light brown to beige, slightly speckled; tail axis yellowish with large dark brown blotches; tail fins with dark patterning, but predominately transparent (in Fig.
All tadpoles were collected from streams in gallery forest and very degraded forest fragments. The description is mainly based on
Long robust tadpole with almost parallel flanks and long, muscular tail (Fig.
The largest specimen at Gosner stage 25, as well as the largest specimen at Gosner stage 36, were the two largest tadpoles in our samples, both reaching about 80 mm total length.
Body light to dark brown, lateral sacs and ventral surfaces lighter, more grayish; fine brown speckling on tail axis, fading from dorsal to ventral parts; fins largely beige transparent, with tiny brown dots along the margin of the last third of dorsal and ventral fin.
Except the large size range of tadpoles all in Gosner stage 25, tadpoles were very uniform in coloration, body and mouth part proportions and characteristics. The tadpole of
All tadpoles were collected from small to medium-sized forest streams. The description is mainly based on
Very long and robust, but still slender tadpole with strong, muscular tail (Fig.
The largest specimen (Gosner stage 25) measured 81 mm total length.
Body and tail including fins plain brown in preservation; ventrum brighter and more grayish; ventral fin brighter and more transparent than dorsal fin (Fig.
All tadpoles were caught in small to medium-sized streams and rivers in low- to middle-altitude forests. The description is mainly based on
Long slender tadpole with slim, muscular tail (Fig.
The largest tadpole (
Body brown, lateral sacs slightly more grayish; tail uniform dark brown at tail base, slightly fading to a fine but dense pattern of lighter brown speckles towards tail tip; fins yellowish brown with a dark reddish brown pattern.
All investigated tadpoles exhibited a similar and complex lateral line-system (LLS), which is usually well developed (Fig.
General arrangement of lateral line-system in tadpoles of the genera
Morphological variability between the investigated tadpoles was relatively minor. We compared various indices (ratios) of body measurements against each other and performed PCAs (e.g. Fig.
Comparison of two morphological characters (body width and body height) against body length in the tadpole species investigated;
The genus
Habitats of
Herein, we describe for the first time the tadpoles of five
Body proportions of
The tadpole of
Tadpoles of
In contrast to the morphological similarity of the herein examined tadpoles, the three genera are markedly different to tadpoles of other arthroleptids. According to
We thank the Cameroonian Ministry of Forestry and Wildlife (MINFOF), and the Guinean Ministère de l’Enseignement Superieur et de la Recherche Scientifique (MESRS) and the Direction Nationale de la Recherche Scientifique et Technique for issuing research, collection and export permits and our field assistants for their help during data collection. We thank D. Blackburn (Florida Museum of Natural History, Gainsville, USA) for a comparison sequence for
Measurements (in mm) of the arthroleptid tadpoles described in this study; given are museum accession numbers, Gosner stage (
Species | Individual | Gosner |
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25 | – | 25.0 | – | 10.7 | 14.2 | 12.5 | 9.2 | 4 | 2.8 | 6.56 | 7.6 | 5.4 | 1.1 | 8.8 | 5.4 | 2.2 | 3.6 | 2.3 | 0.7 | 5 | |
25 | – | 6.8 | – | 2.6 | 3.5 | 3.4 | 2.21 | 1.4 | 1 | 1.7 | 2 | 1.4 | 0.2 | 3.6 | 1.4 | 0.6 | 0.8 | 0.8 | 0.25 | 1.2 | ||
25 | – | 5.3 | – | 2.7 | 3.3 | – | 1.7 | – | – | 1 | 1.1 | 1 | – | – | 0.5 | 0.3 | 0.2 | 0.5 | 0.1 | 0.4 | ||
25 | – | 8.04 | – | 3.74 | 4.66 | 3.8 | 1.93 | 1.4 | 1.1 | 1.78 | 2.3 | 2 | 0.2 | 3.7 | 1.9 | 0.7 | 1.2 | 1 | 0.25 | 1.6 | ||
25 | 20.7 | 7.6 | 13.1 | 3.3 | 4.6 | 3.9 | 1.8 | 1.5 | 1 | 1.6 | 2.7 | 2 | 0.4 | 3.8 | 2.5 | 0.6 | 1.5 | 0.9 | 0.2 | 1.3 | ||
25 | 25.2 | 7.9 | 17.3 | 4.3 | 5.6 | 4.1 | 1.9 | 1.7 | 1.1 | 2 | 3.4 | 2.3 | 0.5 | 4.3 | 2.7 | 0.9 | 1.6 | 1 | 0.2 | 1.5 | ||
36 | – | 24.6 | – | 12 | 13.92 | 11.85 | 9.12 | 4 | 3 | 9 | 9.2 | 6.1 | 1.5 | 11 | 7.9 | 3 | 5 | 3 | 0.8 | 5.1 | ||
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25 | – | 18.71 | – | 7.5 | 9.9 | 8.75 | 5.67 | 3.1 | 2.1 | 4.38 | 6.1 | 4.6 | 0.7 | 9.4 | 4.7 | 1.8 | 3.2 | 2 | 0.7 | 3.5 | |
25 | – | 17.74 | – | 7.08 | 9.8 | 8.61 | 6.5 | 2.5 | 1.8 | 4.59 | 5.4 | 3.9 | 0.8 | 7.6 | 4.2 | 1.7 | 2.8 | 1.9 | 0.5 | 3.4 | ||
25 | 52.8 | 17.3 | 35.5 | 9.3 | 11.6 | – | 7 | 2.8 | 1.8 | 6.3 | 6 | 4.1 | 1.1 | 7.8 | 4.7 | 1.6 | 3.2 | 2.1 | 0.5 | – | ||
36 | 72.1 | 23.7 | 48.4 | 12.7 | 16.1 | 13.3 | 9.2 | 4.8 | 3.2 | 7.1 | 9.6 | 6.5 | 1.4 | 10.5 | 6.5 | 2 | 4.8 | 3 | 0.7 | 5.1 | ||
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25 | – | 14.35 | – | 5.89 | 8.92 | – | 4.29 | – | – | 2.92 | 4.35 | 3.36 | 0.5 | 7 | 3.2 | 0.9 | 2.3 | 1.6 | 0.6 | 3 | |
25 | 58.7 | 21 | 37.7 | 8.7 | 11.9 | 9 | 6.6 | 3.3 | 2.2 | 6.5 | 8.4 | 6.5 | 1.4 | 10.7 | 7.5 | 2.8 | 4.6 | 2.5 | 0.6 | 7.1 | ||
25 | 45.2 | 17.7 | 27.5 | 6.3 | 9.4 | 7.7 | 5.5 | 2.9 | 1.7 | 4.9 | 6.6 | 4.2 | 1.2 | 8.3 | 5.6 | 2 | 3.5 | 2.1 | 0.5 | 3.3 | ||
25 | 74.75 | 25.44 | 49.37 | 10.96 | 13.42 | 9.84 | 6.34 | 4.3 | 3.5 | 5.74 | 7.8 | 6.1 | 1.3 | 12.3 | 5.9 | 1.9 | 4.1 | 2.8 | 1.2 | 5.5 | ||
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25 | 97.38 | 30.86 | 67.01 | 10.77 | 14.93 | 14.9 | 9.73 | 5.45 | 3.99 | 7.86 | 9 | 7.8 | 1.1 | 15 | 7.5 | 2.4 | 5.2 | ||||
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25 | 51.7 | 17.7 | 34 | 7.4 | 9.7 | 9.6 | 5.9 | 3.6 | 2.6 | 4.6 | 7 | 5.4 | 1.6 | 8.1 | 4.8 | 1.2 | 3.7 | ||||
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39 | 62.2 | 22.7 | 39.5 | 6.7 | 15.8 | 8.6 | 5.6 | 2.3 | 1.9 | 5.6 | 9.3 | 3.2 | – | – | 5.5 | 2.1 | 3.5 | 2.5 | 0.5 | 4.9 | |
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25 | – | 22.48 | – | 10.18 | 12.57 | 10.71 | 7.77 | 3.7 | 2.6 | 6.53 | 7.2 | 6.6 | 1.2 | 11 | 5.6 | 1.9 | 3.2 | 2.4 | 0.7 | 4.7 | |
25 | – | 20.19 | – | 7.92 | 11.95 | 8.41 | 6.38 | 2.8 | 2.3 | 5.4 | 5.4 | 5.2 | 1 | 9 | 5 | 1.8 | 3.4 | 2.1 | 0.6 | 4.1 | ||
25 | 60.4 | 20.1 | 40.3 | 7.6 | 10 | 7.4 | 5 | 2.4 | 1.9 | 5 | 7 | 6.5 | 1.1 | 9.9 | 6 | 1.6 | 4.3 | 2.1 | 0.5 | 3.6 | ||
25 | – | 14.65 | – | 6.8 | 9.36 | 6.19 | 5.56 | 2 | 1.5 | 4.62 | 4.3 | 3.7 | – | 6.7 | 3.7 | 1.1 | 2.4 | 1.5 | 0.4 | 3.3 | ||
28 | 73.2 | 23.8 | 49.4 | 12.2 | 12.7 | 12.8 | 8.7 | 4 | 2.5 | 8.4 | 9.1 | 6.7 | 1.2 | 12.2 | 8.5 | 2 | 5.4 | 2.1 | 0.9 | 5 | ||
25 | 52.5 | 18.3 | 34.2 | 7.7 | 9.2 | 8.3 | 5.6 | 3 | 1.9 | 4.9 | 6.1 | 4.5 | 1 | 9 | 5.2 | 2 | 3.2 | 1.8 | 0.6 | 3.4 | ||
25 | – | 11.03 | – | 4.98 | 6.9 | 3.96 | 2.97 | 1.6 | 1.1 | 2.37 | 3.4 | 3 | 0.4 | 5.3 | 2.7 | 0.9 | 1.8 | 1.3 | 0.4 | 2.2 | ||
25 | – | 8.79 | – | 3.81 | 4.91 | 3.67 | 2.35 | 1.5 | 1 | 1.8 | 2.7 | 2.2 | 0.3 | 4.3 | 2.1 | 0.8 | 1.4 | 1 | 0.3 | 1.6 | ||
25 | 34.7 | 11.4 | 23.3 | 5.6 | 6.5 | 4.6 | 2.4 | 1.9 | 1.3 | 2.6 | 4.5 | 3.4 | 0.8 | 5.2 | 3.2 | 0.7 | 2.6 | 1.2 | 0.4 | 2.1 | ||
25 | – | 21.9 | – | 8.4 | 11.1 | – | 5.8 | – | – | 5.8 | 9 | 5.3 | 1.2 | 10 | 2.3 | 2.1 | 5 | 2.4 | 0.4 | 4.1 | ||
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25 | 75.9 | 25.4 | 50.5 | 10.6 | 15.4 | 13.3 | 9 | 4.1 | 3.5 | 7.5 | 9.6 | 5.9 | 1.6 | 10.8 | 6.3 | 1.9 | 4.5 | 3 | 0.6 | 5.9 | |
25 | – | 24.5 | – | 9.9 | 15.2 | 12 | 9 | 4.2 | 3.3 | 7.3 | 9.4 | 5.9 | 1.6 | 10.4 | 5.5 | 1.3 | 4.5 | 2.4 | 0.6 | 5.5 | ||
26 | 80.5 | 24.3 | 56.2 | 9.5 | 13.8 | 10 | 10 | 3 | 1.2 | 9 | 8.9 | 5.6 | 1.7 | 8.9 | 6.2 | 2 | 4.5 | 2.5 | 0.8 | 5 | ||
37 | 80.5 | 28.4 | 52.1 | 10.6 | 17 | 15.1 | 8 | 5 | 3.8 | 7 | 9.6 | 6 | 1.8 | 12 | 6.5 | 1.7 | 5.5 | 3.2 | 0.7 | 5.3 | ||
41 | 77.5 | 24 | 53.5 | 10.6 | 13.3 | 11.5 | 7 | 3.9 | 2.7 | 6 | 9.6 | 5.4 | 1.8 | 11.9 | 6.6 | 2 | 4.3 | 4 | 0.6 | 4.7 | ||
25 | 73.6 | 20.7 | 52.9 | 8.6 | 10.1 | 9 | 7 | 3 | 2.1 | 6.3 | 7.9 | 5.2 | 1.4 | 10.8 | 7 | 2.2 | 4.4 | 2.3 | 0.6 | 4.5 | ||
26 | – | 28.5 | – | 10.8 | 14.8 | 14.4 | 9.3 | 5.1 | 3.9 | 8.6 | 10.2 | 6.4 | 1.4 | 13 | 8.4 | 2.5 | 5 | 3.5 | 0.8 | 5.4 | ||
25 | 95.9 | 27.4 | 68.5 | 10.9 | 15.3 | 12.4 | 9.5 | 4 | 3.1 | 8.6 | 10.2 | 6.5 | 1.6 | 11 | 6.5 | 2 | 5 | 2.5 | 0.8 | 5.3 | ||
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25 | – | 20.01 | – | 8.82 | 11.96 | 9.2 | 6.09 | 2.7 | 2.4 | 5.4 | 6.4 | 5.7 | 0.5 | 10.8 | 5.3 | 1.6 | 3.8 | 2.2 | 0.9 | 4.4 | |
25 | – | 24.91 | – | 11.09 | 13.36 | 11.09 | 9.87 | 3.5 | 3 | 8.11 | 7.9 | 6.6 | 0.4 | 13.2 | 6.4 | 1.5 | 5 | 2.9 | 0.5 | 5.6 | ||
25 | – | 24.27 | – | 10.1 | 13.38 | 10.46 | 7.67 | 3.5 | 3 | 5.9 | 7.6 | 6.9 | 0.8 | 13.5 | 7 | 2.3 | 4.7 | 2.9 | 1.2 | 5.6 | ||
25 | 58.6 | 16.9 | 41.7 | 8 | 9.1 | 7.3 | 5.3 | 2.4 | 2 | 5.1 | 6 | 5.3 | 1.3 | 8.2 | 5.1 | 1.3 | 4.2 | 1.9 | 0.6 | 3.9 | ||
25 | – | 26.24 | – | 10.96 | 13.09 | 12.44 | 8.08 | 3.5 | 2.9 | 6.91 | 8.2 | 7.4 | 0.5 | 20 | 7.9 | 2.2 | 5.7 | 2.9 | 1.2 | 5.8 | ||
25 | – | 8.22 | – | 4.02 | 5.55 | 3.5 | 2.56 | 1 | 0.7 | 1.83 | 3 | 2.5 | 0.5 | 4.9 | 2 | 0.6 | 1.4 | 1.2 | 0.4 | 1.9 | ||
25 | – | 5.31 | – | 1.94 | 2.49 | 2.39 | 1.78 | 1.1 | 0.9 | 1.36 | 2.2 | 2.1 | 0.3 | 3.5 | 1.4 | 0.6 | 0.8 | 0.7 | 0.2 | 1 | ||
25 | 18.6 | 5.9 | 12.7 | 3 | 3.1 | 3 | 1.8 | 1.1 | 0.9 | 1.2 | 1.9 | 1.8 | 0.4 | 2.9 | 1.5 | 0.5 | 1.1 | 0.7 | 0.2 | 1 | ||
25 | – | 7.59 | – | 3.33 | 4.31 | 2.84 | 2.11 | 1.4 | 1 | 1.44 | 3.3 | 2.7 | 0.5 | 4.9 | 2.4 | 0.8 | 1.6 | 1.1 | 0.3 | 1.9 | ||
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27 | – | 30 | – | 11 | 14.3 | 12.1 | 9 | 3.8 | 3 | 8.8 | 10.3 | 9 | (?) 1.5 | 13.8 | 9.9 | 2.4 | 7.9 | 3.2 | 1 | 5.9 | |
28 | 108.1 | 35.8 | 72.3 | 14.9 | 20.9 | 16.2 | 11.8 | 4.5 | 3.1 | 11.3 | 13 | 10.4 | 3 | 16.8 | 11.9 | 3 | 9.1 | 3.6 | 1 | 7.8 | ||
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25 | 18.2 | 6.2 | – | 2.8 | 3.6 | 3.4 | 2.1 | 1.1 | 0.9 | 1.2 | 1.8 | 1.5 | 0.2 | 2.8 | 1 | 0.35 | 0.7 | 0.6 | 0.1 | 0.9 | |
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25 | – | 19.47 | – | 9.09 | 11.98 | 8.67 | 6.15 | 2.9 | 2.2 | 4.95 | 6.1 | 5.4 | 0.9 | 10.1 | 5.6 | 1.6 | 4 | 2.3 | 0.7 | 4.9 | |
25 | – | 13.73 | – | 5.5 | 7.3 | 5.96 | 4.5 | 1.9 | 1.4 | 3.34 | 4.4 | 3.8 | 0.5 | 7.1 | 1.7 | 1.3 | 2.6 | 1.6 | 0.5 | 3 | ||
25 | – | 18.36 | – | 6.97 | 11.67 | 8.42 | 5.25 | 2.8 | 1.9 | 4.33 | 5.5 | 4.8 | 0.5 | 9 | 4.5 | 1.3 | 3.3 | 2.2 | 0.6 | 4.4 | ||
25 | 79.8 | 26.5 | 53.3 | 12.5 | 12.3 | 11.5 | 9.2 | 3.8 | 2.9 | 7.1 | 9.4 | 6.2 | 1.6 | 11.5 | 7.5 | 2 | 5.6 | 2.5 | 0.6 | 5.5 | ||
25 | – | 22.86 | – | 9.68 | 13.49 | 10.81 | 7.32 | 3.6 | 2.5 | 6.37 | 7.4 | 6.5 | 0.5 | 10.4 | 6.6 | 2.2 | 4.6 | 2.4 | 0.8 | 5.3 | ||
25 | – | 19.51 | – | 8.38 | 11.25 | 8.79 | 5.89 | 2.7 | 2 | 5.13 | 6.1 | 5.3 | 0.5 | 9.5 | 4.9 | 1.6 | 3.5 | 2.1 | 0.7 | 4 | ||
25 | – | 13.68 | – | 5.48 | 7.23 | 5.85 | 3.93 | 1.8 | 1.3 | 2.92 | 4.2 | 3.8 | 0.4 | 7.1 | 3.9 | 1.3 | 2.5 | 1.7 | 0.5 | 3.1 | ||
25 | – | 17.24 | – | 7.37 | 10.63 | 7.37 | 5.1 | 2.6 | 1.9 | 4.3 | 5.1 | 4.3 | 0.4 | 7.3 | 4.9 | 1.7 | 3.2 | 1.9 | 0.6 | 4 | ||
25 | – | 19.3 | – | 8.08 | 11.62 | 8.16 | 5.98 | 3.1 | 2.2 | 5.2 | 6.4 | 5.2 | 0.5 | 10 | 5.7 | 1.8 | 3.9 | 2.3 | 0.7 | 5.5 | ||
25 | – | 19.61 | – | 7.95 | 11 | 8.96 | 6.4 | 3 | 2.2 | 5.3 | 6.5 | 5 | 0.4 | 9.7 | 4.4 | 1.5 | 3.1 | 2.2 | 0.7 | 4.2 | ||
25 | – | 18.75 | – | 7.94 | 10.14 | 8 | 6.14 | 2.5 | 2.2 | 4.44 | 5.6 | 4.9 | 0.6 | 9.2 | 5.3 | 1.9 | 3.4 | 2.1 | 0.7 | 4.4 | ||
25 | 61.5 | 19.1 | 42.4 | 7.6 | 9.4 | 8 | 5.2 | 2.6 | 1.8 | 4.3 | 6.5 | 5.4 | 0.9 | 9 | 6.5 | 1.8 | 4.5 | 2 | 0.6 | 4 | ||
25 | – | 26.18 | – | 10.81 | 13.57 | 11.1 | 8.43 | 4.2 | 3.1 | 7.75 | 8.3 | 7.1 | 1 | 13.8 | 8.2 | 2.8 | 5.4 | 3 | 1 | 5.9 | ||
25 | – | 14.42 | – | 6.27 | 8.78 | 6.26 | 4.39 | 2.2 | 1.5 | 3.65 | 4.7 | 3.9 | 0.4 | 7.2 | 3.9 | 1.3 | 2.8 | 1.7 | 0.6 | 3.3 | ||
25 | – | 12.01 | – | 5.69 | 7.29 | 5.65 | 4.01 | 1.7 | 1.3 | 3.3 | 4 | 3.5 | 0.5 | 6.3 | 3.1 | 1 | 2.2 | 1.3 | 0.4 | 3.3 | ||
25 | – | 13.65 | – | 6.55 | 8.72 | 5.98 | 4.28 | 1.8 | 1.2 | 3 | 4.7 | 3.5 | 0.5 | 6.8 | 3.7 | 1.1 | 2.6 | 1.2 | 0.5 | 2.8 | ||
25 | – | 12.84 | – | 5.65 | 7.33 | 4.26 | 4.66 | 1.7 | 1.4 | 2.79 | 3.9 | 3.4 | 0.4 | 6.4 | 3.4 | 1.1 | 2.4 | 1.5 | 0.5 | 3.2 | ||
25 | – | 23.28 | – | 9.57 | 12.03 | 9.78 | 6.46 | 3.3 | 2.6 | 5.43 | 7 | 6.2 | 0.7 | 10.7 | 6.3 | 1.9 | 4.4 | 2.1 | 0.8 | 4.9 | ||
39 | 74.2 | 24 | 50.2 | 9.9 | 12.1 | 11.4 | 5.5 | 4.5 | 2.7 | 5.1 | 8.5 | 7 | 2 | 11.4 | 8 | 1.9 | 6 | 2.9 | 0.6 | 6 | ||
25 | 38.8 | 13.5 | 25.3 | 6 | 8.5 | 6.5 | 4 | 2.1 | 1.3 | 3.8 | 5.5 | 4.1 | 0.5 | 6 | 4.4 | 1.1 | 3.6 | 1.3 | 0.4 | 3.2 | ||
36 | 77 | 27.5 | 49.5 | 10.5 | 12 | 10.5 | 7 | 3.6 | 2.7 | 6.2 | 9.5 | 7.5 | 1.6 | 13.2 | 8.7 | 2.3 | 6.1 | 2.6 | 0.6 | 5.1 | ||
36 | 79.2 | 25.6 | 53.6 | 9.9 | 12.5 | 10.6 | 7 | 3.2 | 2.3 | 6.8 | 9.4 | 7.5 | 2 | 12.4 | 8.5 | 2.1 | 6 | 2.5 | 0.7 | 5.1 | ||
26 | – | 23.4 | – | 10.2 | 12.7 | 11.1 | 7.6 | 3.4 | 2.5 | 7.4 | 8.5 | 7 | 1.6 | 12 | 8 | 2.4 | 5.6 | 2.5 | 0.6 | 5.9 | ||
26 | 76.4 | 23.6 | 52.8 | 11.1 | 12.3 | 11.5 | 8.5 | 4 | 2.6 | 7.8 | 9.5 | 7.9 | 1.5 | 11.5 | 8.5 | 2 | 6.5 | 2.5 | 0.5 | 6.1 | ||
27 | – | 24.1 | – | 9.4 | 12.1 | 8.6 | 6.5 | 3.4 | 2.1 | 5.6 | 8.2 | 6.5 | 1.1 | 11 | 7.4 | 2.1 | 5.1 | 2.4 | 0.6 | 4.5 | ||
26 | 74.2 | 23.2 | 51 | 9.5 | 12.3 | 9 | 6.5 | 2.9 | 2.6 | 5.9 | 8.1 | 6.9 | 1.1 | 11 | 7.9 | 2.1 | 5.3 | 2.5 | 0.7 | 5.1 | ||
25 | 52.9 | 17.6 | 35.3 | 6.5 | 9.6 | 7 | 5.5 | 2 | 1.6 | 4.4 | 6.4 | 4.9 | 1 | 8.1 | 6 | 1.6 | 4.3 | 2 | 0.5 | 3.3 | ||
|
25 | 81.2 | 31.6 | 49.6 | 8 | 12.6 | 10.6 | 6.8 | 3 | 2.3 | 7.1 | 8.1 | 6.4 | 1.1 | 10.5 | 7.8 | 2.5 | 5.4 | 2 | 0.6 | 6.5 | |
25 | – | 22.9 | – | 6.6 | 9 | 8.5 | 5 | 3 | 2.4 | 5 | 5.6 | 5 | 0.9 | 9 | 6.5 | 2 | 4.5 | 1.6 | 0.5 | 5.3 | ||
26 | 69.1 | 25.6 | 43.5 | 8.6 | 10.1 | 9.1 | 5.2 | 2.5 | 1.8 | 5.2 | 7 | 6 | 1 | 10.5 | 8.5 | 2.5 | 5.2 | 1.9 | 0.5 | 6.4 | ||
25 | – | 27.2 | – | 7.1 | 10.4 | 9.5 | 5.5 | 3.5 | 2.7 | 5.2 | 6.9 | 5.9 | 1.4 | 11 | 8.5 | 3.2 | 5.3 | 1.6 | 0.5 | 6.6 | ||
25 | – | 21.4 | – | 6.7 | 8.6 | 7.9 | 5.8 | – | – | 4.8 | 5.8 | 4.5 | 1.1 | 8 | 6 | 2 | 3.9 | 1.1 | 0.4 | 4.4 | ||
|
25 | – | 6.3 | – | 1.9 | 3.5 | – | 1.2 | – | – | 1.2 | 2.3 | 1.5 | 0.5 | 3.5 | 1.9 | 0.5 | 1.2 | 0.8 | 0.1 | 1.1 | |
25 | 20.4 | 6.7 | 13.7 | 2 | 3.7 | 3.1 | 1.4 | 1 | 0.6 | 1.3 | 2.5 | 1.8 | 0.6 | 3.5 | 2.2 | 0.6 | 1.2 | 0.6 | 0.1 | 1.2 | ||
25 | – | 10.4 | – | 4.6 | 5.4 | – | 3.2 | – | – | 2.9 | 3.5 | 2.4 | – | – | 2.4 | 1.3 | 2 | 1.2 | 0.1 | 2.1 | ||
41 | 62.2 | 19 | 43.2 | 8.3 | 10.9 | 8.5 | 5.9 | 3.3 | 2.2 | 4.1 | 7.2 | 2.5 | 0.9 | 9 | 4.3 | 0.5 | 2.7 | 2.4 | 0.2 | 3.6 | ||
25 | 37.5 | 11.8 | 25.7 | 4.5 | 6.6 | 5.1 | 3.8 | 1.9 | 1.4 | 3 | 4.5 | 2.6 | 1.1 | 6.5 | 3.3 | 1.1 | 2.1 | 1.4 | 0.2 | 2 | ||
39 | 57.7 | 18.7 | 39 | 9.2 | 12.2 | 9 | 7 | 3.3 | 2.3 | 4.1 | 7.5 | 4.2 | 1.6 | 9.8 | 5.2 | 1.6 | 3.5 | 2 | 0.2 | 3.8 | ||
25 | – | 12.9 | – | 5.3 | 6.8 | – | 4 | – | – | 3.5 | 5 | 3.2 | 0.8 | 7 | 4.2 | 1.6 | 2.6 | 1.5 | 0.2 | 2.4 | ||
25 | 47.5 | 15.5 | 32 | 5.9 | 8.6 | 6.8 | 4.5 | 2.7 | 1.6 | 3.2 | 5.8 | 3.9 | 0.9 | 7.6 | 4 | 1.2 | 2.7 | 1.7 | 0.2 | 3 | ||
25 | 48.3 | 16.1 | 32.2 | 6.2 | 8.9 | 7.5 | 5.2 | 2.9 | 1.7 | 3.4 | 6.4 | 3.5 | 0.9 | 8 | 4.1 | 1.2 | 2.8 | 1.9 | 0.2 | 2.9 | ||
38 | 55.3 | 18.7 | 36.6 | 8.1 | 10.3 | 9 | 6.8 | 3.2 | 1.9 | 5 | 7 | 3.8 | 1.6 | 9 | 5.1 | 1.6 | 3.6 | 2.3 | 0.2 | 3.5 | ||
25 | 36.6 | 11.9 | 24.7 | 5.2 | 6.8 | 5.7 | 4 | 1.9 | 1.3 | 2.8 | 4.5 | 2.7 | 1 | 6.7 | 3.5 | 1.2 | 2.4 | 1.6 | 0.1 | 2.4 |