a) Bayesian inference tree based on 529 bp of the mitochondrial 16S. Asterisks denote Bayesian posterior probabilities values: one asterisk enclosed in parentheses, ≥ 90%; one asterisk, ≥ 98%; two asterisks, ≥ 99–100%. b) Haplotype network reconstruction for the phased alleles of the nuclear POMC gene fragment in P. laevis from Betampona, Marovato and Anivorano Est, P. alluaudi from Tsitongambarika and Andohahela, P. sp. Ca01 from Ambatolahy and Rhombophryne sp. from Torotorofotsy.