Distribution of P. biseriatus, P. tanganicus and P. cornusafricae sp. nov. in the Horn of Africa. Localities sampled for the genetic analysis are shown along with sample codes. The background layer shows shaded relief and water bodies and drainages and was made with Natural Earth (www.naturalearthdata.com). The species range maps were derived from Roll et al. (2017, 2021), Spawls et al. (2023), and the findings presented in this study.