Research Article |
Corresponding author: Maxim V. Vinarski ( radix.vinarski@gmail.com ) Academic editor: Matthias Glaubrecht
© 2016 Maxim V. Vinarski.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Vinarski MV (2016) Annotated type catalogue of lymnaeid snails (Mollusca, Gastropoda) in the collection of the Natural History Museum, Berlin. Zoosystematics and Evolution 92(1): 131-152. https://doi.org/10.3897/zse.92.8107
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The article deals with examination of the type materials of sixty-one species and variety of lymnaeid snails (Mollusca: Gastropoda: Lymnaeidae) housed in molluscan collection of the
Pond snails, taxonomy, nomenclature, type series, history of malacology
The malacological collection of the Berlin
In this article, I present the results of my examination of the type series of species belonging to the family Lymnaeidae Rafinesque, 1815 housed in
This study is a part of my recent project devoted to identification and publication of the Lymnaeidae types from European repositories. I worked with
Abbreviations of shell dimensions are as follows. SH – shell height, SW – shell width, SpH – spire height, BWH – body whorl height, AH – aperture height, AW – aperture width, WN – whorls number. All dimensions in the taxa accounts are given in millimeters.
BMNH – British
Limnaea
alfredi
Limnaea
tenisoni
var. β
Alfredi
Lymnoёa
alfredi
Limnaea
alfredi
Limnaea
alfredi
Lymnaea
truncatula
The lectotype is housed in the Museum of New Zealand (Te Papa Tongarewa) under accession number M 125077 (see
New Zealand, Southern Island, Governors Bush, Hooker Valley, Mount Cook Hermitage (fide
Type specimens of Lymnaeidae (
Limnaea (Gulnaria) ovata var. amnicola
Limnaea
ovata
var.
amnicola
It is known that Westerlund often distributed parts of the type series of taxa described by him among several European museums (
Ronneby and Kristianstad, Sweden (
An obvious junior synonym of Radix (Peregriana) balthica (L., 1758).
Limnaeus
amygdalum
Limnaeus
amygdalum
Limnaea acuminata var. amygdalum von Martens, 1881: 76, pl. 14, figs 7, 8.
Limnaea
acuminata
var.
amygdalum
Limnaea
acuminata
f.
typica
Lymnaea
auricularia
race
rufescens
Lymnaea (Pseudosuccinea) acuminata Subba Rao 1989: 126, figs 254–265, 272.
Two samples of L. amygdalum from the Ganges River in
India, the Ganges River.
Radix (Radix) rufescens (Gray, 1822).
Limnaea javanica var. angustior von Martens, 1881: 88, pl. 16, fig. 8.
Limnaea javanica var. angustior von Martens, 1897a: 4.
Limnaea javanica var. angustior Kilias, 1961: 162.
49 syntypes collected in Makassar (Celebes Island, Indonesia) and kept under No. 8136. The largest of these shells reaches 26.2 mm height.
Indonesia: Java Island (Batavia and Tji-surupan), Celebes Island (Makassar). leg. von Martens.
Cerasina luteola (Lamarck, 1822).
Limnaea
stagnalis
var.
baltica
Limnaea
stagnalis
var.
baltica
Limnaea
stagnalis
f.
baltica
Baltic Sea, east shores of the Gotland Island (Östergarn, Legraf, Haugröne and Fårosund).
Lymnaea (Lymnaea) stagnalis. This variety represents a dwarf morph of the great pond snail inhabiting the Baltic Sea (
Limnaea
brevispira
von
Lymnaea
brevispira
Lymnaea
brevispira
A single specimen is kept in
WN 2.50; SH 10.9; SW 9.5; SpH 1.2; BWH 10.4; AH 9.6; SW 6.8.
Indonesia, Sumatra Island, Manindjau Lake.
This is valid species with unclear generic position. It may belong to either genus Austropeplea Cotton, 1942 or to the genus Bullastra Bergh, 1901.
Limnaeus
cerasum
Limnaea
acuminata
var.
cerasum
Lymnaea
luteola
There are three samples of L. cerasum from the Ganges River in
India, the Ganges River.
A junior synonym of Cerasina luteola.
Limnaea
columella
var.
championi
Limnaea
columella
var.
championi
Two syntypes collected in Bugaba (South Panama) by Champion and kept in
Panama (southern), Bugaba.
Pseudosuccinea columella (Say, 1817).
Limnaea
auricularia
var.
coreana
von
Lymnaea
auricularia
var.
coreana
Lymnaea (Radix) coreana
Lymnaea (Radix) coreana
Lymnaea (Radix) coreana
11 syntypes in two samples: No. 38440 (seven syntypes) and No. 55594 (4 syntypes), leg. Gottsche (without date).
Korea, “Changjin, Prov. Hangyöngdo”.
Radix (Radix) coreana.
See Table
Measurements of shells of syntypes of some lymnaeid species described by von Martens (
Character / index | Limnaea auricularia var. coreana |
Limnaea eversa No. 34822 (n = 10) |
Limnaeus patulus No. 8044 (n = 25) |
|
---|---|---|---|---|
No. 38440 (n = 7) | No. 55594 (n = 4) | |||
Whorls number |
3.5 – 3.75 3.62 ± 0.11 |
3.50 ± 0.00 |
3.87 – 4.50 4.08 ± 0.21 |
5.00 – 5.50 5.20 ± 0.13 |
SH, mm |
24.8 – 29.4 27.9 ± 1.5 |
27.5 – 31.8 28.9 ± 2.0 |
14.4 – 17.3 15.7 ± 1.0 |
34.6 – 44.4 39.0 ± 2.9 |
SW, mm |
18.2 – 23.6 21.3 ± 1.7 |
22.1 – 26.5 23.7 ± 2.0 |
10.4 – 12.3 11.2 ± 0.6 |
16.2 – 24.6 18.9 ± 2.3 |
SpH, mm |
3.4 – 5.4 4.4 ± 0.7 |
3.3 – 7.1 4.7 ± 1.7 |
4.3 – 5.9 4.9 ± 0.5 |
9.8 – 16.6 13.0 ± 1.7 |
BWH, mm |
22.8 – 27.3 25.6 ± 1.5 |
22.1 – 29.9 25.4 ± 3.5 |
12.8 – 15.2 13.7 ± 0.8 |
29.4 – 39.3 33.7 ± 2.8 |
AH, mm |
20.2 – 27.4 24.5 ± 2.4 |
24.8 – 26.7 25.9 ± 0.8 |
11.2 – 13.4 12.2 ± 0.8 |
24.8 – 33.8 28.2 ± 2.6 |
AW, mm | 14.6 – 19.8 17.8 ± 1.6 |
18.2 – 19.2 18.8 ± 0.4 |
8.2 – 10.0 8.9 ± 0.8 |
11.8 – 19.1 14.5 ± 2.2 |
SW/SH |
0.73 – 0.80 0.76 ± 0.03 |
0.79 – 0.85 0.82 ± 0.03 |
0.68 – 0.77 0.72 ± 0.02 |
0.42 – 0.55 0.48 ± 0.03 |
SpH/SH |
0.14 – 0.20 0.16 ± 0.02 |
0.12 – 0.22 0.16 ± 0.04 |
0.27 – 0.35 0.31 ± 0.02 |
0.24 – 0.39 0.48 ± 0.04 |
BWH/SH |
0.90 – 0.93 0.92 ± 0.01 |
0.79 – 0.94 0.88 ± 0.07 |
0.84 – 0.89 0.87 ± 0.02 |
0.82 – 0.89 0.86 ± 0.02 |
AH/SH |
0.80 – 0.95 0.89 ± 0.05 |
0.84 – 0.93 0.90 ± 0.04 |
0.73 – 0.81 0.78 ± 0.03 |
0.66 – 0.80 0.72 ± 0.03 |
AW/AH |
0.70 – 0.79 0.73 ± 0.03 |
0.70 – 0.76 0.73 ± 0.02 |
0.64 – 0.78 0.74 ± 0.05 |
0.49 – 0.64 0.51 ± 0.04 |
Type specimens of Lymnaeidae (
Limnaea
lagotis
var.
costulata
von
Limnaea
lagotis
var.
costulata
Limnaea lagotis f. costulata Annandale & Rao 1925: 153, fig. 7.
Limnaea
auricularia
var.
lagotis
f.
costulata
Lymnaea
costulata
Radix
lagotis
var.
costulata
I could not recognize the syntype(s) of this variety in
Uzbekistan, Tashkent City, leg. A.P. Fedchenko.
Radix (Peregriana) lagotis (Schrank, 1803) or Radix (Radix) sp.
In 1897, von
Limnaeus
cubensis
Limnaeus
cubensis
Limnaea cubensis von Martens 1899: 378.
Galba (Galba) cubensis
Lymnaea
cubensis
Limnaeus
cubensis
Fossaria (Bakerilymnaea) cubensis
Fossaria
cubensis
Two syntypes kept under No. 101522. leg. Pfeiffer.
“Cuba”, without precise location.
(1) WN 5.25; SH 8.1; SW 4.4; SpH 4.5; BWH 5.9; AH 3.8; SW 3.1; (2) WN 5.25; SH 7.6; SW 4.1; SpH 4.1; BWH 5.2; AH 3.6; SW 2.7.
Galba (Bakerilymnaea) cubensis. Some authors place this species into the genus Fossaria Westerlund, 1885.
Amphipeplea
cumingiana
Amphipeplea
cumingi
Lymnaea
cumingiana
Two probable syntypes from Luzon Island kept under No. 109771. leg. Dunker ex coll. Pfeiffer. Another probable syntype from the same island is under No. 109772. The largest of these specimens is 26.1 mm height. The labels bear no information about the nomenclatorial status of the shells, and their identification as probable syntypes may be questioned. The species name on the labels is spelled as “Amphipeplea cumingi” (see Remark below).
Philippines, island of Luzon, Naga, province of South Camerines. leg. Cuming.
Bullastra cumingiana.
Originally,
Limnaea
amygdalum
var.
cycacea
Lymnaea
auricularia
race
rufescens
Five syntypes kept under No. 109768.
India, the Ganges River.
Radix (Radix) rufescens (Gray, 1822).
Limnaea
elmeteitensis
Lymnaea
elmeteitensis
Limnaea
elmeteitensis
Lymnaea
natalensis
Three syntypes of L. elmeteitensis kept under No. 47554. The largest of them is 21.2 mm height. The rest of the type series is, probably, in BMNH.
Kenya, lakes Baringo and Elmeteita.
Radix (Radix) natalensis (Krauss, 1848).
Stagnicola
elrodi
USA, Montana, west shore Flathead Lake, 13 1/2 miles north of Poison.
WN 5.25; SH 16.3; SW 8.6; SpH 7.6; BWH 13.2; AH 10.2; SW 5.8.
Limnaea
ovata
var.
eversa
von
Limnaea
auricularia
var.
eversa
Lymnaea eversa Starobogatov & Streletzkaya 1967: 231, fig. 18.
The lectotype of L. ovata var. eversa (see Fig.
Northern Mongolia, Eter River near Dzha-Dzassyk Monastery. leg. Potanin, 1877.
Radix (Peregriana) balthica (L., 1758). Most of the syntypes correspond to the species Lymnaea (Peregriana) intermedia Lamarck, 1822 sensu
See Table
Limnaeus
natalensis
var.
exsertus
von
Limnaeus
natalensis
var.
exsertus
Limnaea
exserta
von
Lymnaea
natalensis
Lymnaea
natalensis
A single shell (the syntype) is kept under No. 8586. This shell has 9.4 mm height.
Ethiopia (Abyssinia), Aiz Zaba spring near Zasaga.
Radix (Radix) natalensis (Krauss, 1848).
Amphipeplea
ampulla
var.
globosa
Amphipeplea
ampulla
var.
globosa
Amphipeplea
ampulla
var.
globosa
Simlimnaea
tomentosa
Amphipeplea
ampulla
var.
globosa
Lymnaea
tomentosa
tomentosa
There are three syntypes in
New Zealand, Southern Island, Governors Bush, Hooker Valley.
Austropeplea tomentosa (Pfeiffer, 1855) [fide
Limnaea
acuminata
var.
gracilior
von
Limnaea
acuminata
var.
gracilior
Lymnaea
auricularia
race
rufescens
Lymnaea (Pseudosuccinea) acuminata Subba Rao 1989: 126, figs 254–265, 272.
There is a single specimen (the syntype) of L. acuminata var. gracilior in
India, Bengal (without precise location).
Radix (Radix) rufescens.
Limnaea
gutta
Limnaea gutta Kilias, 1967: 339.
There is a single specimen (syntype) of L. gutta in
Italy, Lombardy (from the Villa and Villa’s work title).
The syntype of L. gutta may be identified as a juvenile R. (Peregriana) sp. Possibly, this shell belongs to R. balthica or R. ampla (Hartmann, 1821).
Type specimens of Lymnaeidae (
Limnaea
humerosa
von
Lymnaea
natalensis
Limnaea
humerosa
Lymnaea
natalensis
The lectotype (
Uganda, Mengo, in an artificial pond leg. Stuhlmann (05.01.1891) – locality of the lectotype. von
Radix (Radix) natalensis.
Stagnicola
impedita
Stagnicola
impedita
Originally, the type series consisted of four specimens (the holotype and three paratypes) housed in the Geological Department of the Stanford University under accession number 5776 (
USA, Utah, near Logan, Cash Co.
Limnaeus
javanicus
var.
intumescens
von
Limnaea
javanica
var.
intumescens
von
Limnaea
javanica
var.
intumescens
von
Limnaea
javanica
var.
intumescens
Three shells of syntypes collected in Surabaya (Java Island, Indonesia) and kept under No. 8140. The largest of these shells reaches 26.6 mm height.
Indonesia: Java Island (Surabaya, Passuruan, Rogodjampi). leg. von Martens and Zollinger.
Cerasina luteola.
Limnaea
kempi
Lymnaea
natalensis
Limnaea
kempi
Lymnaea
natalensis
A single (subadult) specimen, the syntype, is kept in
East Africa, Victoria Lake.
Radix (Radix) natalensis.
Limnaea
limosa
var.
ovata
f.
margaritacea
Limnaea
lagotis
var.
margaritacea
Limnaea
lagotis
var.
margaritacea
Limnaea
lagotis
var.
margaritacea
Two syntypes are kept in
Ronneby, Sweden.
Most probably, Radix (R.) auricularia.
Limnaea
stagnalis
var.
westerlundi
f.
nereni
Lymnaea
stagnalis
Limnaea
stagnalis
var.
westerlundi
f.
nereni
Lymnaea
stagnalis
Two syntypes are kept in
Skeninge, Sweden.
Lymnaea (Lymnaea) stagnalis.
Limnaea
nitidella
von
Limnaea
nitidella
The lectotype (designated by
Ecuador, region of Chorrera de Agoyan (von
WN 3.75; SH 7.8; SW 5.1; SpH 2.9; BWH 6.5; AH 5.0; SW 3.3.
Uncertain.
Limnaeus
nucleus
Limnaea
acuminata
var.
nucleus
von
Limnaeus
nucleus
Limnaea
acuminata
var.
nucleus
Limnaea
luteola
f.
ovalis
Ten syntypes are kept in
India, the Ganges River.
A junior synonym of Cerasina luteola.
Limnaea
nyansae
von
Lymnaea
nyansae
Limnaea
nyansae
Lymnaea
natalensis
16 shells (syntypes) from the type locality are kept in
The western shore of the Victoria Lake, near Bukoba and Towalio.
Radix (Radix) natalensis.
Limnaeus
javanicus
var.
obesus
von
Limnaea
javanica
var.
obesa
von
Limnaeus javanicus var. obesus Kilias, 1961: 164, fig. 5.
14 shells of syntypes collected in Indonesia and kept under No. 8124. The largest of these shells reaches 27.0 mm height.
Indonesia: Java Island (Telaga, Patengan). leg. Baron von Richthofen.
Cerasina luteola.
Type specimens of Lymnaeidae (
Limnaeus
obliquatus
von
Limnaea
auricularia
var.
obliquata
Radix
auricularia
var.
obliquata
Lymnaea (Radix) obliquata
Two syntypes (
“Im Landsee Issyk-Kul am Nordabhang des Thienschan, 4691´ Pariser Fuss über der Meere, 43° N.B“ = Kyrgyzstan, nothern shore of the Issyk-Kul’ Lake.
Radix (Radix) obliquata.
(1) WN 3.75; SH 27.1; SW 25.2; SpH 7.3; BWH 26.3; AH 23.3; SW 16.8; (2) WN 4.12; SH 28.8; SW 25.8; SpH 6.8; BWH 27.4; AH 22.4; SW 17.6.
The type series has been overlooked by
Limnaeus
patulus
Limnaeus
patulus
Limnaea
acuminata
var.
patula
Limnaea
acuminata
var.
patula
Lymnaea
auricularia
race
rufescens
Lymnaea acuminata f. patula Subba Rao 1989: 127, figs 256, 257.
The syntypes of L. patulus in
India, the Ganges River.
Radix (Radix) rufescens.
See Table
Limnaeus
pervius
von
Limnaea
pervia
von
Limnaeus
pervia
Lymnaea
pervia
Galba
pervia
Limnaeus
pervius
The lectotype (designated by
Northern China, Chi-foo (= Tschifu). leg. Schottmüller.
Orientogalba hookeri (Reeve, 1850).
Limnaea (Gulnaria) pettiti
Lymnaea
pettiti
Limnaea
pettiti
East China, “near Chefoo, Shantung” (= Shandong Province).
Possibly, a synonym of Radix (Radix) plicatula (Benson, 1842).
Limnaea
javanica
var.
porrecta
von
Limnaea
javanica
var.
porrecta
von
Limnaea
javanica
var.
porrecta
There are 14 shells of this variety collected from the type locality in
Timor Island, Kupang. leg. E. von Martens, December 1862.
Cerasina luteola (Lamarck, 1822).
Limnaeus
prunum
Limnaea
acuminata
var.
prunum
Limnaeus
prunum
12 syntypes of L. prunum in
India, the Ganges River (
Cerasina luteola (Lamarck, 1822).
Limnaea (Gulnaria) ovata var. sericina
Limnaea
ovata
var.
sericina
Five syntypes are kept in
Ronneby, Sweden.
Most probably, Radix (P.) balthica.
The syntypes (
Limnaea (Gulnaria) shantungensis
Lymnaea
shantungensis
Limnaea
shantungensis
East China, “Shantung, Wei Hai Wei” (= Shandong Province).
Orientogalba viridis (Quoy & Gaimard, 1834).
Limnaea
ovata
var.
solidior
von
Northwestern China, Dzungaria, Ulungur River. leg. A. Regel, 1879.
Possibly, Radix (Peregriana) lagotis.
Limnaea
lagotis
var.
solidissima
Limnaea
lagotis
var.
solidissima
Limnaea
lagotis
var.
solidissima
East India, Himalaya Mts. (without precise location).
WN 4.00; SH 17.4; SW 13.2; SpH 6.6; BWH 15.3; AH 12.9; SW 8.8.
Radix (Peregriana) lagotis.
Limnaea
solidulus
Limnaea solidulus Kilias, 1967: 340.
There is a single specimen (syntype) of L. solidulus in
Italy, Brescia (from the syntype label).
The syntype of L. solidulus may be identified as a juvenile R. (Peregriana) balthica = Lymnaea intermedia Lamarck, 1822 sensu
Limnaea
amygdalum
var.
straminea
Lymnaea
auricularia
race
rufescens
31 syntypes kept in
India, the Ganges River.
Radix (Radix) rufescens (Gray, 1822).
Limnaeus
subulatus
Limnaeus
subulatus
11 shells from the type locality in three samples: no. 4613 and two without numbers. One of these shells (see fig. 45) is separated and marked as belonging to the type collection (a syntype). The rest of specimens are not formally labeled as syntypes but probably also origin from the type series.
Mexico, in Zimapan and Lake of Mexico (
Ladislavella (Walterlymnaea) elodes (Say, 1821).
WN 7.75; SH 32.2; SW 9.6; SpH 22.1; BWH 18.3; AH 11.8; SW 6.6.
Not discussed by
Type specimens of Lymnaeidae (
Limnaea
javanica
var.
subteres
von
Limnaea
javanica
var.
subteres
von
Limnaea
javanica
var.
subteres
There is a single shell of this variety in
Indonesia, Sumatra Island, Palembang. leg. E. von Martens.
Possibly, a synonym of Radix (Radix) rubiginosa (see
Limnaeus
sulcatulus
Limnaea
acuminata
var.
sulcatula
Lymnaea
auricularia
race
rufescens
Three samples in
India, the Ganges River.
Radix (Radix) rufescens.
Limnaea
tugrina
Limnaea
luteola
f.
ovalis
Lymnaea
luteola
A single shell (syntype) is housed in
Ceylon (without precise location).
Cerasina luteola (Lamarck, 1822).
Limnaea undussumae von Martens 1897: 135, pl. I, fig. 18; pl. VI, figs 2, 5.
Lymnaea
undusumae
(sic!)
Lymnaea
natalensis
The lectotype (designated by
Undussuma, in a brook beyond the Tamaro. leg. Stuhlmann, 1891.
Radix (Radix) natalensis.
Bullastra
velutinoides
Lymnaea
cumingiana
The only shell (syntype) is kept in
Bullastra cumingiana.
Limnaea (Gulnaria) whartoni
Lymnaea
whartoni
Limnaea
whartoni
East China, “Liu Shi Tao, north-east promontory of Shantung”.
Possibly, Radix plicatula.
Limnaea
lagotis
var.
yarkandensis
From near Sásak Taka (
Radix (Peregriana) lagotis.
Limnaeus coarctatus Dunker (? in MS).
Cerasina luteola.
The species has not been mentioned in the most inclusive handbooks on lymnaeid taxonomy (
Limnaeus compactus “Ziegler” (? in MS).
Two shells collected in the Danube River in Austria are housed in
An obvious synonym of R. auricularia.
The species has not been mentioned in the most inclusive handbooks on lymnaeid taxonomy (
Limnaea cornea “Ziegler” (? in MS).
Three shells collected in Carniolia (= Kraina, a historical region of Slovenia) and kept in
Radix peregra = R. labiata sensu
L. cornea is absent in most taxonomic works devoted to overview of the European continental malacofauna, including those dealing with the species names attributed to Ziegler (
Limnaea elgonensis Preston (? in MS).
A single specimen kept under No. 62871. Its shell height is 12.0 mm.
Uganda, Mt. Elgon. leg. Preston.
Probably, a junior synonym of Radix natalensis.
I could not find the original description of this species in Preston’s works devoted to new taxa of African land and freshwater mollusks (
Limnaea fernanensis Preston (? in MS).
A single shell is kept under No. 63775. This shell has 19.2 mm height.
«British East Africa» (probably Kenya), Fort Fernan.
Radix (Radix) natalensis.
See remark for L. elgonensis above.
Limnaeus nebulosus Dunker (? in MS).
Two shells (labelled as ‘types’) collected in the Antilles (without precise locality) and kept in
Most probably, a synonym of Galba cubensis.
This species name is absent in subsequent comprehensive works devoted to overview the New World lymnaeid fauna (
Limnaea acuminata var. nevilli von Martens (? in MS).
A single shell (SH = 32.3 mm) collected in Bengal (without precise location) by Lamare Piquot is kept in
Radix (Radix) rufescens.
I could not find this variety name in von Martens publications devoted to Indian freshwater snails (von
Limnaeus nigricans “Ziegler” (? in MS).
A single shell, 10.8 mm shell height, collected in Neuwaldegg, Austria, is kept in
An obvious synonym of Radix (Peregriana) peregra (O.F. Müller, 1774) auct. = Radix labiata (Rossmässler, 1835) sensu
See comment to Limnaeus compactus “Ziegler” above.
Limnaeus nitens “Ziegler” (? in MS).
Two shells collected in Vienna, Austria, are kept in
Possibly, a synonym of Radix (Peregriana) balthica.
See comment to Limnaeus compactus “Ziegler” above.
Limnaea
opaca
Limnaeus
pereger
var.
opacus
von
Limnaea
peregra
var.
opaca
Two shells collected in Carniolia and kept in
Possibly a synonym of Radix (Peregriana) peregra auct. = Radix labiata sensu
Though a few authors of the 19th century used this taxon name (
Amphipeplea pfeifferiana Dunker (? in MS).
Five shells collected somewhere in New Holland (= mainland Australia) are kept in
Possibly, a synonym of Austropeplea lessoni (Deshayes, 1830) .
Like other species names attributed to Dunker and listed in this supplement, L. pfeifferiana seems to be a member of the group of so-called “manuscript names”, whose original descriptions have been not found in the literature.
Limnaeus splendens Dunker (? in MS).
Six shells from China (without precise location) kept under accession number 109769. The largest of them is 19.2 mm height.
Possibly, a synonym of Radix plicatula.
See remark to Amphipeplea pfeifferiana above.
I wish to express my sincerest thanks to Dr. Matthias Glaubrecht (Hamburg, Centrum für Naturkunde) and Christine Zorn (Berlin,